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Table 5.4. Effects of otter trawls on mud substrate habitat: summary of published studies. (S = statistically significant; citations in bold print are peer-reviewed publications.)

No. Reference Location Depth Sediment Effects Recovery Approach
1 Ball et al. 2000 Irish Sea 75 m Sandy silt Reduced infaunal and epifaunal richness, diversity, number of species, and individuals in fishing ground compared to wreck site, but no obvious effects on macrofauna 24 hr after trawling.   Experimental trawling in heavily fished prawn fishing ground, unfished area near a shipwreck used as control.
2 Brylinsky et al. 1994 Bay of Fundy, Nova Scotia, Canada Intertidal Silt and coarse sand overlain with silt Door tracks in sediment, rollers compressed sediment; S decrease in nematodes and benthic diatoms in door tracks, no effects on larger infaunal organisms (mostly polychaetes). Furrows visible 2-7 mo; nematodes recovered in 1-1.5 mo, diatoms in about 1-3 mo. Four trawling experiments (repeated tows during a single day) at two locations in a trawled area, effects evaluated for 1.5-4 mo.
3 DeAlteris et al. 1999 Narragansett Bay, Rhode Island, USA 14 m Mud Doors produced tracks 5-10 cm deep and adjacent berm 10-20 cm high. No changes in hand dug trenches for >60 days. Diver observations.
4 Drabsch et al. 2001 Gulf of St. Vincent, South Australia 20 m Fine silt Trawl door tracks, smoothing of topographic features; S decrease in total infaunal abundance and one group of polychaetes, damaged epifauna.   Experimental trawling (two tows per unit of area in 1 day) in area with no trawling for 15 yrs (one site); effects evaluated after 1 wk.
5 Frid et al. 1999 Northeast England (North Sea) 80 m Silt-clay S increase in total number of individuals in taxa predicted to increase at high fishing effort and number of errant polychaetes; no effect of increasing effort on total number of individuals expected to decrease, but S decline in sea urchins.   Related changes in benthic fauna in a heavily trawled location to low, high, and moderate fishing activity and to changes in phytoplankton production over 27 yr.
6 Hansson et al. 2000 Fjord on the west coast of Sweden 75-90 m Clay Abundance of 61% infaunal species negatively affected and S reductions in abundance of brittle stars during last 5 mo of disturbance period; S reductions in total biomass at 3 of 3 trawled sites and 1 of 3 control sites, and in number of individuals at 2 of 3 trawled sites and 1 of 3 control sites; abundance of polychaetes, amphipods, and mollusks not affected.   Experimental trawling for 1 yr (two tows per wk, twenty-four tows per unit of area) in area closed to fishing for 6 yr (three treatment and three control sites); effects evaluated during last 5 mo of experiment.
7 Mayer et al. 1991 Maine coast, USA 20 m Mud Dispersal of fine surface sediment; doors made furrows several cm deep; some planing of surface features, but no plowing of bottom or burial of surface sediments.   Experimental trawling (single tow); examined immediate effects on sediment composition and food value to sediment depth of 18 cm.
8 Pilskaln et al. 1998 Gulf of Maine, USA 250 m Mud Greater abundance of suspended infaunal polychaetes in more heavily trawled area.   Deployed sediment traps in fishing grounds 25-35 m above substrate.
9 Sanchez et al. 2000 Coast of Spain, Mediterranean Sea 30-40 m Mud Door tracks in sediment; no change in number of infaunal individuals or taxa, or in abundance of individual taxa; no changes in community structure. Door tracks still clearly visible after 150 hr. Experimental trawling in trawled area at two sites swept once and twice in a single day; effects evaluated after 24, 72, 102, and 150 hr.
10 Sparks-McConkey and Watling 2001 Penobscot Bay, Maine, USA 60 m Mud S decline in porosity, increased food value, and increased chlorophyll production of surface sediments; S reductions in number of infaunal individuals and species, species diversity, and abundances of 6 polychaete and bivalve species, S increase in nemerteans. All geochemical sediment properties and all but one polychaete/bivalve species recovered within 3.5 mo, nemerteans still more abundant after 5 mo. Experimental trawling (four tows in 1 day) in untrawled area; pre-trawl sampling of sediments and infauna for a year; recovery monitored for 5 mo.
11 Tuck et al. 1998 West coast of Scotland 30-35 m Fine silt Tracks in sediment, increased bottom roughness; no effect on sediment characteristics; S increase in number of infaunal species at end of 16 mo disturbance period and during 18 mo recovery period; no change in biomass or number of individuals at end of recovery period; S increase in polychaetes, S decrease in bivalves; mixed results of analyses of community structure, S reduction in diversity during first 22 mo. Door tracks still evident after 18 mo; bottom roughness recovered after 6 mo; nearly complete recovery of infaunal community within 12 mo, complete after 18 mo.  Experimental trawling for 1 day/mo (one and a half tows per unit of area) for 16 mo in area closed to fishing for >25 years; recovery monitored after 6, 12, and 18 mo.

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